OPTIMUS PRIME
Optronix was a data clerk and a pupil of the sage Alpha Trion. He grew tired of collecting data and decided to join the Autobot Guard. He became best friends with the Elite Guard commander Tronion and the two of them fought crime together. Their friendship came to a sudden end, however, when Tronion came across an ancient relic, which corrupted him with power and granted him the name Megatron. He then went on to begin an uprising in the fallen city Kaon, starting the Decepticon faction.
When the Great War, Optronix abandoned his name, seeing himself as someone different than who he was now, and took on the name Optimus. He became a commander in the Autobot military and became Autobot leader Zeta Prime's top lieutenants. After Zeta was killed in battle by Decepticon forces, the High Council chose Optimus to be the next Prime, giving him the Matrix of Leadership.
Optimus Prime's first act as leader was to create an Ark that would send him and a crew of Autobots to find energy for their home world. As the Ark took off, Megatron's henchmen spotted the ship and the Decepticon leader took off after them. The two factions later ended up on a crash course for the planet Earth. There they laid dormant for four million years, until they were awoken by an volcanic eruption and their war continued.
Many events would happen throughout the war, including the reawakening of Atha Prime, the return of Nova Prime, and the arrival of Unicron. By the time the Chaos Bringer was defeated, Optimus, having seen enough action, passed on the Matrix to Rodimus Prime, while he joined the reformed High Council in the late Alpha Trion place.
As time went on, however, Optimus noticed that Rodimus was unsure of his position as leader and decided to step down from the council so that he could mentor Rodimus. A short time later, the Cybertronian Empire invaded the Milky Way Galaxy, with Cybertron being its primary target. Optimus and Rodimus led the Autobots against the invaders, who nearly managed to conquer their world had it not been for Rodimus, who sacraficed his life to wipe out the Empire.
For a brief moment, everything seemed to beat peace, until Megatron, who had been thought dead, returned and challenged Optimus to one final duel. Optimus obliged and charged towards his former comrade. Their duel lasted long, until Optimus gathered up his strength and courage and killed Megatron once and for all.
In battle's aftermath, Optimus decided it was time for him to retire and he passed on leadership to Omega Prime. Optimus then traveled to a highly populated planet with two suns, which he would stay for the next three years.
Optimus quickly came out of retirement when he was informed that his son, Orion Pax (or Optimus Minor), was being hunted by Decepticons. He returned to Earth to face another former friend of his: Nemesis Prime.
Optimus quickly defeated Nemesis and then went out to save Orion from the Fallen. He entered a duel with the ancient being, along with two other Decepticons, and valiantly risked his life to buy his son time to escape. Despite using all his might, Optimus was sneaked up by the Fallen and stabbed through the chest.
His sacrifice was not in vain however, and his son managed to end the war against the Decepticons. After the Matrix was recovered from Omega Prime's deceased body, it was buried with Optimus, for he was the last true Prime in the eyes of the Autobots.
MEGATRON / GALVATRON
Tronion was a commander in the Autobot Elite Guard during Sentinel Prime's age, and was considered by the population of Iacon to be one of its greatest crime fighters. When Optronix joined the Guard, he and Tronion became best friends, along with Optronix's friends Magnus, Dion, and Ariel. Their friendship came to an unfortunate end however, when he was corrupted by an ancient relic containing the Fallen's essence and became Megatron, leader of the evil Decepticons.
Throughout the following war, Megatron was feared by Autobot and Decepticon alike. During a surprise attack on Iacon, Megatron confronted the Autobot leader Zeta Prime and killed him with his bare hands.
Later, when the Autobot ship Ark took off, Megatron took a crew of Decepticons with him on board the Nemesis and pursued Optimus Prime and his Autobots. The two factions ended up on Earth, where, four million years later, their war would continue.
Megatron and Optimus would frequently clash during the Atha Prime and Expansion conflicts. After the defeat of Nova Prime and the Fallen, Megatron was imprisoned in the Stockade. Months later, he was freed by the Heralds of Unicron and was brought before the Chaos Bringer himself. Megatron reluctantly agreed to become Unicron's herald in exchange for a new, powerful form. He was then reformatted into Galvatron!
Galvatron returned to Cybertron, where his Decepticons were divided by civil war. He managed to end the conflict and take control of the Decepticons. He was then ordered by his master to retrieve the energies of Vector Sigma. Once doing so, Galvatron returned to Unicron.
During the Chaos Bringer's attack on Cybertron, Galvatron confronted the young Autobot Hot Rod. He was shortly defeated just as Hot Rod used the Matrix of Leadership to destroy Unicron.
Some time later, Galvatron returned, being neither Decepticon nor Herald and set out to find Cybertron's core. He then merged himself with the core and essentially became Cybertron itself. He chased after the rebelling Autobots to the planet Gaia, where he transformed his planet form into a giant robot and prepared to destroy Gaia. Before he could do so, the Phantom Violen Jygar intervened and took over Galvatron's body. After Rodimus defeated the Phantom, Galvatron was reverted to his original form, which was now heavily mutilated and was left for dead on an unidentified planet.
He was later found by Necros and Nova Prime and was resurrected into Megatron once again. He took back command over the Decepticons and returned to Cybertron, where he challenged Optimus Prime to a final duel. The two leaders fought for quite some time, until he was at Optimus' mercy. For a brief second, Megatron felt remorse for his actions, but the feeling faded as Optimus brought his Energon Axe down and ended Megatron's life.
In the years that followed, many Decepticons would attempt to achieve victory in his honor, but none of them succeeded in doing so. By the time the Cybertron Alliance was formed, Megatron was regarded with both honor and disdain by the Decepticons.
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